Interesting notice, Greg! It reminds me of an old experiment taught at school: if you cross-pollinate a pea with green, smooth seeds and a variety with yellow, rough seeds, you'll get plants producing green rough seeds, and yellow ones with a smooth surface (and of course some of the hybrids will produce similar seeds to their parents' seeds). So the color of cotyledons is inheritable from both parents.
It would be interesting to check how it works. One could cross a citrus variety with green cotyledons and, for example, a trifoliate orange. Then plant the green and white seeds separately and check if the white seeds produced trifoliate seedlings.