Hello,
Photosynthesis... I had a long discussion about photosynthesis with a citrus fellow here in Germany, and we fought a strong blade battle, which his sword as sharp as mine!
Beside all the chemical things happaning in the photosynthesis process, mostly for energy resorbtion things and energy absorbtion control, the main things happen physically
So, Millet and Skeeter,
Citrus is a common plant of the C3 typus, unlike like corn,
Zea mays, which is C4 typus, has a saturisation level of solar radiation. About one thrid sunlight is the saturisation level in
Citrus depending on Severtson (University of Florida). All above is not needed, and further resorbition of light is controlled by the mentioned chemical processesm, but mainly by control of evaporation and light resorbtion surface.
So shortly:
Citrus controls the photosynthesis by evaporation and in low day temperature more water is used in photosyntheses for gluccose production, than for evaporation. As more sunlight reaches the leaf surface, as higher the evaporation get's and lower the absorbtion of Carbondioxide will get, simply to maximize the evaporation of water. Evaporation has a cooling effect on the leaf lamina, preventing the leaf form getting to hot by solar radiation, because radiation not used by the photosyntheses process will by absorbed by the leaf cells, and transfered into heat (enrgy transfer from radiation energy to heat energy) This heat is the source of all sunburn!
And that's why plants with cold roots in direct sunlight, usually in spring outdoors suffer more from sunburn, as other plants. Because the cool spring nights and the cool morning temperatures inhibit high water uptake rates by the roots, and so sunliught hits the leaf unshaded with the full energy. Evaporation levels increase rapidly, but the roots won't be able to deliver the full evaporation level of water need to cool the leaf lamina, so local overheating of leaf areas will happen, causing sunburn.
But beside this process,
Citrus has two other, additional ways of preventing to much solar radiation to the leaf surface.
First is the oil film on the lamina surface. The oil glances in the leaf will dispense oils to the leaf surface. We all know this, because the smell of these essential oils is in full sunlight much higher, than in morning hours. This oil will generate a film on the leaf lamina and may biologists consider this shiny film as an additional protection agains solar radiation, like a liquid sunscreen.
The shiny oil film will reflect light, and thus prevent sunlight from reaching the lamina surface or penetrate the cellular structure of the leaf, preventing the heat up process inside the plants leaf.
Next protection is a leaf curl. As one can observe:
Citrus leaves have more silvery leaf downside. Simpl go to your plant and turn one leaf upside down and watch the downside, you will surely remark the silvery glance...
As solar radiation gets higher and higher, evaporation increases. If evaporation levels reach excess, and leaf damage may occur, because of more water is need for evaporation to prevent overheating as the whole system is able to evaporate, (Sorry, I do not know how to tell you correctly...) the plant curls the leaf. The leaf margins roll from side to the midvein, so shading the leaf lamina by itself and exposing the leaf downside to the sun, with it's shiny, bright color. So it seems, that the leaf downside has a much higher reflective ability than the shiny oil film, so the combination of shading, reduced exposed leaf surface and the bright leaf downside seems to reduce very effective the radiation stress to the leaf surface, and prevents overheating and excess evaporation in high temperature, high radiation circumstances.
Well, sorry for my bad english, but I hope you will understand...