There was some work done back in the late 60s or early 70s, with mature groves of oranges on rough lemon roots in central Florida. Groves planted on opposite sides of US highway 27, in Polk County -- a 4-lane US highway, with a wide median, and wide area off to each side. Light, well-drained sandy soils. So the groves were likely at least 200 feet apart (I'm sorry I don't have the written reference to this -- I'm remembering it from my intro to citrus class that I took at UF). There was evidence of roots intermingling out in the middle of the median, allowing transfer of nematodes. So at least in that case -- mature, rough-lemon-rooted trees, a radius of over 100 feet was possible. That doesn't mean the bulk of the roots were out there, and we still recommend irrigation and fertilizer under the canopy drip line; but at least some roots can go very much farther.